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What does it take to be a scientific pioneer? 

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To reframe and popularise evolutionary theory? 

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To reveal a new material, and win science's most coveted prize? 

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To discover one of palaeontology's elusive missing links? 

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Is the key to brilliance talent, ego or just plain good luck? 

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What makes a beautiful scientific mind? 

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Professor Jenny Clack is a world-renowned palaeontologist, 

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who solved one of the greatest riddles in the history of life on our planet. 

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One of the big questions that people had been trying to answer was, 

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how do you get from an animal that lives in the water and has fins, 

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to an animal that walks on land and has limbs with fingers and toes? 

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In the late 1980s, she found and described the fossil Acanthostega, 

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an ancient creature which offered new evidence 

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of how fish evolved legs and made the transition onto land. 

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Her talent for seeing what others had missed rewrote the textbooks 

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and led to global recognition for her work. 

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She's the great pioneer. 

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She's the one who led the way 

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and opened up this area that we others have come into. 

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We do talk about the Clack theory of the origin of tetrapods. 

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It does sometimes do to think about things that you take for granted. 

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What if it's not this way, but it's the other way? 

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But becoming a pioneer was by no means inevitable. 

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She's not a person with sharp elbows. That's not it at all. 

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It was quite clear she didn't have a great deal of confidence in herself. 

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Clack had to find the courage to take on 

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the stifling academic etiquette that had hampered research for decades. 

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The field had become moribund, and now it's been completely unlocked. 

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Answering one of evolution's greatest mysteries would mean 

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travelling to almost the ends of the earth for fossil evidence. 

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I was actually terrified, but discovery of those materials 

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was probably the most exciting thing after falling in love. 

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Just what does it take to turn accepted thinking on its head, 

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and make the palaeontological discovery of a generation? 

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Jenny Clack has made a life's work out of trying to find traces of a world 

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hundreds of millions of years ago, before even the dinosaurs. 

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Her passion is for the very first creatures that emerged from the seas 

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to conquer the Earth. 

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For me, trying to imagine what these animals were like, 

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it's a sense of mystery, where the animals are totally different 

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from anything that we have today. 

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It's trying to just imagine what they were like and what they were 

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doing, and what life was like for them, and just wanting to be there. 

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But the evidence is elusive. 

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With only scant fossilised remains to go on, 

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seeing into the rocks poses a creative challenge. 

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That has a lot to do with wanting to see something else 

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in the specimen that nobody's ever seen before, 

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and being able to see a little bit more about it. 

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It all goes into this sort of imagination 

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of what the animal would have been like. 

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Jenny's fascination for this era, known as the Devonian, 

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has fuelled her imagination since early childhood. 

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What sparked my interest in palaeontology 

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was an illustrated book called Prehistoric Animals, 

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and it had sections on the earliest part of the fossil record, 

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and it was always those earlier sections that really intrigued me, 

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and I remember flicking through this book, starting at the beginning, 

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and listening to the slow movement of Shostakovich's Fifth Symphony, 

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and if you do that, the music fits the pictures perfectly. 

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And if you know that piece of music, 

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that kind of sums up how I imagine the Devonian to be. 

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The story of the colonisation of the land by the creatures that 

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emerged from the swamps and pools of the Devonian Period 

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is one of the most fascinating episodes in the history of life. 

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But this mysterious world was a long way from the Manchester suburb 

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where Jenny grew up. 

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It wasn't a particularly special household. 

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A simple terraced house, fronting onto a road. 

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My parents were not at all academic. 

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In fact, they were very poorly educated, 

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and I don't remember the house being full of books, for example, 

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apart from the ones I borrowed from the library. 

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But we had the great advantage that, 

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across the road from where I lived, was a pond, 

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and in that pond were sticklebacks and tadpoles and all sorts 

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of wonderful things, and I used to bring home sticklebacks and newts. 

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My mother hated newts, but she let me keep them anyway. 

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And that was just a wonderful resource. 

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I was an only child, so I would go there regularly by myself. 

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Intrigued by the primitive-looking animals she found on her doorstep, 

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Jenny soon discovered there was another way 

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she could connect with creatures from the distant past. 

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Through fossils. 

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The remains of these animals will be down in the rocks, 

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well below the surface of the Earth, 

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but where cliffs like this have broken away, 

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then the rocks are exposed, and we can see where the animals' remains are. 

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By the time I was 11 or 12, 

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I'd heard of this great Victorian fossil hunter called Mary Anning, 

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who lived and worked at Lyme Regis, 

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and found some wonderful specimens like the fish lizards, 

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ichthyosaurs, and I decided that I wanted to be the new Mary Anning. 

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So we duly went to Lyme Regis. 

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While most children were playing on the beach, 

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Jenny was engrossed by the fossils. 

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I didn't find any ichthyosaurs, but we did find lots of ammonites, 

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and I thought to myself, "Wow, that sounds wonderful. 

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"I really want to do that." 

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Jenny had found her life's passion, 

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but for a girl growing up in the 1950s, 

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palaeontology wasn't seen as an obvious choice. 

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'Science plays an important part in the curriculum, these days. 

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'But girls who may be among the leading biologists must also 

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'be able to use their hands, and that's a useful experience 

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'when it comes to shopping in the future.' 

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When I was in my secondary school, 

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I actually felt very much of an also-ran. 

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I don't think I was considered anything special. 

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Certainly, it was never suggested that I would take the Oxbridge exams. 

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When it came to A-levels, they weren't as good as expected, 

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and I actually got into my fourth choice, Newcastle upon Tyne, 

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chosen really on the list at all because, one, it had a palaeontology option, 

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and two, it had a Gilbert and Sullivan society. 

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Don't let's be downhearted! 

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There's a silver lining to every cloud. 

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Certainly! Let's be perfectly happy! 

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By all means! Let's thoroughly enjoy ourselves! 

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It's absurd to cry. 

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Quite ridiculous! 

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Jenny's fourth choice turned out to be a lucky one. 

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The palaeontology option was thriving at Newcastle, 

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under the leadership of the forward-thinking Dr Alec Panchen. 

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'At Newcastle University Department of Zoology, 

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'Dr Alec Panchen talks about the difficulty of deciding 

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'how the first reptiles may have emerged from amphibian ancestors.' 

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If one looks at a reptile, such as a lizard, 

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and then an amphibian, such as a newt or a salamander, 

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the differences between the two aren't so obvious. 

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Dr Panchen's lab was one of the few places in the world 

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looking at the origin of tetrapods, 

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the broad term used for all four-limbed vertebrates, 

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from amphibians to mammals. 

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He was studying the earliest known tetrapods, 

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from over 300 million years ago, 

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precisely the era that had fired Jenny's childhood imagination. 

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But Dr Panchen set a high bar. 

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Alec Panchen was a very meticulous worker. 

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His papers have immense detail. 

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His drawings, beautiful reconstructions of skulls, 

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incredibly detailed. 

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Quite intimidatingly, for his students. 

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We all felt we had to try and draw like that 

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or we would be failures, and it certainly pulled us up a lot. 

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Although keen to study with Panchen, 

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Jenny found it hard to attract his attention. 

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I actually found him quite difficult to talk to. 

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He's very, very reserved. 

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When you get two people who are quite reserved together, 

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the conversation doesn't always flow freely, shall we say. 

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Panchen's elite group of PhD students seemed out of reach. 

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Well, she was very reserved, and in that sense, 

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maybe she didn't feel confident enough, 

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but there was no indication that she had talents 

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that would get her where it finally did. 

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When I got to my final year, I would have liked to do a PhD 

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in palaeontology, 

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but I was also told that Panchen didn't take anybody on 

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who hadn't got a first, 

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and I thought I was unlikely to get a first. 

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And, in fact, I didn't. 

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And so, I sort of gave up on that idea. 

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Instead of applying to join Dr Panchen's exclusive circle 

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of research graduates, Jenny opted for a less challenging job 

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in a regional museum in Birmingham. 

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My first job was a display technician. 

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Well, I wasn't altogether happy as a museum education person, 

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because I'm not that comfortable with children, 

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and of course a lot of the job was working with children. 

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I don't really know how to deal with them. 

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And she might have stayed in that job without the encouragement 

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that came from a chance meeting. 

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When I was working in Birmingham, I got interested in motorcycles. 

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And while I was at motorcycle club, I was part of a folk group, 

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and this guy was doing a floor spot, singing by himself with a guitar, 

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and I don't know what brought this subject up, 

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I don't know why he mentioned it, but he said the word dimetrodon. 

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She heard the magic word and pricked up her ears, and it really 

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was just a matter of time before we actually met from then on. 

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"What is this guy doing mentioning dimetrodon?" 

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Pretty soon we got to know each other. 

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So, yes, we never looked back. 

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Jenny's future husband Rob didn't only share her passion for bikes. 

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Soon the couple were hitting the road as fossil hunters. 

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We certainly would take our holidays on the back of a bike, 

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going up to Scotland to look for fossils. 

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And Rob would prove to be a vital support 

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at times when Jenny was uncertain how to progress. 

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It was quite clear, in those days, that she didn't have a great deal of confidence in herself. 

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But, looking back on it, that was simply a lack of experience. 

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With Rob's encouragement, Jenny kept nurturing her passion for fossils, 

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and it wasn't long before she spotted an opportunity. 

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My boss at the time knew that I wasn't entirely content in that job, 

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so she suggested that I do some private study, 

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because the local authority actually allowed staff to take off 

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three weeks a year to do some private study, 

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and so I took a chance. 

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Jenny knew that Bradford City Museum owned a fossil 

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that had been discovered in a local coal mine. 

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It was of a 300 million-year-old creature called Pholiderpeton, 

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thought to be one of the first tetrapods to venture onto land. 

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Jenny's old university tutor, Alec Panchen, had tried without success to get his hands on it. 

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Many years before, they had put it on display 

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embedded in a block of concrete, 

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and they would never take it out of the concrete for Alec, 

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and he was always very upset that this was the one specimen 

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he had never been allowed to work on. 

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Jenny hatched an idea that she thought just might impress him. 

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Using her museum connections, she promptly went and borrowed it, so she had it. 

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She recognised the power of having a good hand of cards 

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when you negotiate something. 

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Her hunch proved right. 

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Intrigued, Dr Panchen invited her to join him in Newcastle to inspect the specimen. 

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But it wasn't an easy fossil to get to grips with. 

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I remember a colleague of mine having a look at the specimen, 

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and he's used to dealing with roadkills, but he said, 

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"My Goodness, that's a mess!" 

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That was his reaction to the specimen. 

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But it turned out to be much more interesting 

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than you might have expected. 

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Despite having little time to impress her old tutor, 

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Jenny's sharp eye began to reveal itself. 

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I was preparing it, and looking at the bits that hadn't been exposed to the surface before, 

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and I found this peculiar chunky bone, 

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and I showed this to Alec, and I remember his precise words. 

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He said, "Well, I'm damned. It's a braincase." 

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And then, a few seconds later, "There could be a PhD in this. 

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"I could probably get a grant for you, if you're interested." 

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Can a duck swim? 

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Jenny promptly quit her museum job 

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and headed to Newcastle to begin a new life of academic research. 

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The Bradford fossil Pholiderpeton would form the basis of years of painstaking examination. 

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'Here in the Department of Zoology at Newcastle University in England, 

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'a research student is working on the fossil remains of one early amphibian.' 

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This animal lived 300 million years ago, and one can only be 

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fascinated and curious about what life was like in those days. 

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I'm, in fact, the first person to restudy this animal since 1926. 

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'She first of all cleans away parts of the matrix surrounding 

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'the fossil using a dental mallet, then further cleaning up 

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'by blasting with a jet of sodium bicarbonate particles.' 

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You're exposing things that have never been seen by anybody before. 

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It's compulsive. Your sense of time just disappears. 

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And unless somebody comes and knocks on the door and says, 

244
00:16:49,040 --> 00:16:51,520
"Time for a cup of tea," you could just stay there. 

245
00:16:55,440 --> 00:16:58,240
When you go to sleep at night, and you close your eyes, 

246
00:16:58,240 --> 00:16:59,720
you can see it in front of you. 

247
00:17:01,200 --> 00:17:02,800
It's a bit like watching a movie. 

248
00:17:05,480 --> 00:17:07,920
If you prepare your own material, 

249
00:17:07,920 --> 00:17:11,960
it does give you an intimate view of what you've got. 

250
00:17:11,960 --> 00:17:16,960
And this Bradford fossil was a rather strange specimen. 

251
00:17:16,960 --> 00:17:19,800
It had, probably underwater, 

252
00:17:19,800 --> 00:17:24,120
crawled inside a hollow tree trunk and died, 

253
00:17:24,120 --> 00:17:28,200
and then decomposed, and you knew it was all one animal, 

254
00:17:28,200 --> 00:17:31,600
and you knew that you had all the bits. 

255
00:17:31,600 --> 00:17:35,320
If only you could take it apart and put them together the right way. 

256
00:17:35,320 --> 00:17:39,480
As she put the pieces together, Jenny made a startling discovery. 

257
00:17:43,120 --> 00:17:45,000
It was one Friday evening. 

258
00:17:45,000 --> 00:17:46,760
I was working on the airbrasive, 

259
00:17:46,760 --> 00:17:51,080
and there was a little bone right at the tip of the specimen, 

260
00:17:51,080 --> 00:17:57,400
rather apart from all the other bones, so I sat and I prepared it. 

261
00:17:57,400 --> 00:17:59,040
So, this thing is about yay long. 

262
00:18:03,400 --> 00:18:05,240
Like nothing I'd ever seen before. 

263
00:18:06,480 --> 00:18:08,760
And I thought, "Now, wait a minute. 

264
00:18:08,760 --> 00:18:12,560
"The only bones that I can think of which have got holes 

265
00:18:12,560 --> 00:18:16,120
"through like that that aren't vertebrae 

266
00:18:16,120 --> 00:18:21,840
"would be one of the braincase bones at the back called the exoccipital, 

267
00:18:21,840 --> 00:18:24,240
"or a stapes." 

268
00:18:25,440 --> 00:18:30,840
My mind actually couldn't believe it to start with. 

269
00:18:30,840 --> 00:18:34,200
The stapes is a tiny bone in the ear of modern vertebrates. 

270
00:18:34,200 --> 00:18:37,120
Its ability to vibrate is critical to hearing. 

271
00:18:38,640 --> 00:18:41,840
No-one had ever discovered a stapes in an early tetrapod before, 

272
00:18:41,840 --> 00:18:46,760
as the bone was thought to be too small to survive fossilisation. 

273
00:18:46,760 --> 00:18:49,680
But there was something strange about this stapes. 

274
00:18:50,680 --> 00:18:53,360
What the stapes showed that I found 

275
00:18:53,360 --> 00:18:56,720
was that it was completely the wrong shape, 

276
00:18:56,720 --> 00:19:02,800
and it was fixed in a way that meant it couldn't transmit vibrations. 

277
00:19:02,800 --> 00:19:04,560
It wasn't free to vibrate. 

278
00:19:05,720 --> 00:19:08,120
So something odd was going on. 

279
00:19:08,120 --> 00:19:11,760
A stapes that was too big and solid to vibrate 

280
00:19:11,760 --> 00:19:14,160
would be useless for hearing in air. 

281
00:19:14,160 --> 00:19:16,480
But the prevailing theory 

282
00:19:16,480 --> 00:19:19,200
was that early land-dwelling tetrapods must be able to hear. 

283
00:19:20,520 --> 00:19:23,640
Jenny wondered if there was another way of looking at it. 

284
00:19:25,760 --> 00:19:30,480
It does sometimes do to think about things that you take for granted 

285
00:19:30,480 --> 00:19:31,960
and turn them on their heads. 

286
00:19:33,400 --> 00:19:37,880
What if it's not this way, but it's the other way? 

287
00:19:37,880 --> 00:19:42,400
The assumption had been that as fish evolved into tetrapods, 

288
00:19:42,400 --> 00:19:44,760
a bone in the jaw had rapidly transformed 

289
00:19:44,760 --> 00:19:47,480
into the tiny stapes that could vibrate. 

290
00:19:49,560 --> 00:19:52,960
Jenny's discovery suggested that this transformation had not 

291
00:19:52,960 --> 00:19:56,480
occurred as early as previously thought. 

292
00:19:56,480 --> 00:20:01,880
So, the stapes that I found is a step along the way into making 

293
00:20:01,880 --> 00:20:05,960
a modern ear in something like modern reptiles. 

294
00:20:05,960 --> 00:20:08,080
But it doesn't do it straight away. 

295
00:20:09,320 --> 00:20:12,960
Jenny wondered whether the bone she'd identified was in fact 

296
00:20:12,960 --> 00:20:18,200
evidence of a whole sense, hearing, in the process of evolving. 

297
00:20:18,200 --> 00:20:20,960
What if these animals couldn't yet hear? 

298
00:20:23,440 --> 00:20:24,880
Think about it like this. 

299
00:20:25,920 --> 00:20:28,320
What was there for it to listen to? 

300
00:20:28,320 --> 00:20:35,000
There weren't any creaking insects, there weren't any birds, 

301
00:20:35,000 --> 00:20:37,240
no animals were making noises. 

302
00:20:37,240 --> 00:20:40,840
The only sounds would have been the wind in the trees, 

303
00:20:40,840 --> 00:20:43,440
leaves blowing about, that kind of thing. 

304
00:20:43,440 --> 00:20:47,960
She began to trawl through decades of academic descriptions 

305
00:20:47,960 --> 00:20:51,040
of the ears of early tetrapods, 

306
00:20:51,040 --> 00:20:53,640
and soon realised that many of them did have stapes. 

307
00:20:53,640 --> 00:20:56,840
Jenny had spotted something that had been right under peoples' noses 

308
00:20:56,840 --> 00:21:01,800
all along, but simply dismissed as an unimportant scrap of bone. 

309
00:21:01,800 --> 00:21:08,000
Alec Panchen had been working on another related animal for some time. 

310
00:21:08,000 --> 00:21:10,800
There was a stapes, only Alec hadn't described it, 

311
00:21:10,800 --> 00:21:15,560
and Jenny pointed it out to him, you know, there, and it looks the same, 

312
00:21:15,560 --> 00:21:20,840
and he admitted that he had passed over it in embarrassed silence. 

313
00:21:24,560 --> 00:21:28,400
Jenny's work on stapes updated the textbooks. 

314
00:21:28,400 --> 00:21:31,600
Her unique way of seeing was yielding results. 

315
00:21:31,600 --> 00:21:36,640
One of the interesting things about it is to show 

316
00:21:36,640 --> 00:21:40,480
how what you see is governed by what you expect to see. 

317
00:21:41,840 --> 00:21:44,520
And that happens all the time, actually. 

318
00:21:44,520 --> 00:21:48,920
You don't always see what's there, because you've got a certain predisposition. 

319
00:21:48,920 --> 00:21:53,680
It certainly had been excellent training for her, 

320
00:21:53,680 --> 00:21:57,040
and it was clear that this was somebody who was capable of 

321
00:21:57,040 --> 00:22:00,440
changing the way we thought about some of these animals. 

322
00:22:00,440 --> 00:22:03,200
Well, it was at that point that I thought perhaps there were 

323
00:22:03,200 --> 00:22:06,080
other things that people have always taken for granted 

324
00:22:06,080 --> 00:22:08,640
that aren't the way we thought they were, and yes, 

325
00:22:08,640 --> 00:22:12,160
that was a lesson that I've continued to apply. 

326
00:22:14,280 --> 00:22:17,120
That certainly stood me in good stead. 

327
00:22:20,880 --> 00:22:22,680
Jenny was beginning to get noticed. 

328
00:22:23,920 --> 00:22:25,800
She landed a prestigious job 

329
00:22:25,800 --> 00:22:29,640
as a curator at Cambridge University's Museum of Zoology. 

330
00:22:32,800 --> 00:22:36,840
I was quite amazed when I got a job in Cambridge because, 

331
00:22:36,840 --> 00:22:40,560
not having been considered as a candidate for Oxbridge at school, 

332
00:22:40,560 --> 00:22:43,480
there's no way I was going to get into Cambridge. 

333
00:22:43,480 --> 00:22:48,880
I sort of felt that I'd got in by the back door, 

334
00:22:48,880 --> 00:22:51,440
via the museum connection. 

335
00:22:51,440 --> 00:22:55,360
In those days, many of the staff at Cambridge 

336
00:22:55,360 --> 00:23:01,320
still looked down on anyone who was "redbrick", and I think Jenny 

337
00:23:01,320 --> 00:23:05,040
had a lot more problems from being a redbrick than being a woman. 

338
00:23:06,920 --> 00:23:10,640
And certainly, several of them would always refer to her as Mrs Clack, not Dr Clack. 

339
00:23:11,960 --> 00:23:15,480
Aware that critical eyes were upon her, 

340
00:23:15,480 --> 00:23:17,800
and with the freedom now to pursue her own research, 

341
00:23:17,800 --> 00:23:20,960
Jenny chose to tackle one of the biggest mysteries 

342
00:23:20,960 --> 00:23:22,560
in our evolutionary story. 

343
00:23:24,720 --> 00:23:28,840
One that had taxed some of the greatest minds in palaeontology 

344
00:23:28,840 --> 00:23:30,160
for decades. 

345
00:23:32,880 --> 00:23:35,880
One of the big questions that people had been trying to answer, 

346
00:23:35,880 --> 00:23:38,360
in various ways, 

347
00:23:38,360 --> 00:23:44,040
was how do you get from an animal that lives in the water and has fins, 

348
00:23:44,040 --> 00:23:48,200
to an animal that walks on land and has limbs with fingers and toes? 

349
00:23:49,640 --> 00:23:54,240
And the evidence was simply absent. 

350
00:23:54,240 --> 00:23:56,760
Many of us would say it's more radical than anything that's 

351
00:23:56,760 --> 00:24:01,880
taken place on land, and that the origin of reptiles, 

352
00:24:01,880 --> 00:24:05,200
the origin of birds, the origin of mammals are all, really, 

353
00:24:05,200 --> 00:24:08,840
less radical than this fish-to-tetrapod transition. 

354
00:24:12,280 --> 00:24:15,760
It was thought that this important transition 

355
00:24:15,760 --> 00:24:21,040
from fish to vertebrates with limbs occurred during the Devonian era, 

356
00:24:21,040 --> 00:24:23,720
which began 400 million years ago. 

357
00:24:24,960 --> 00:24:28,480
But there was precious little evidence of how it had happened. 

358
00:24:29,560 --> 00:24:32,920
Imagination is very important in any science. 

359
00:24:32,920 --> 00:24:37,000
You have to be able to think about things that you can't see, 

360
00:24:37,000 --> 00:24:42,200
but exist in your head, and for a palaeontologist, 

361
00:24:42,200 --> 00:24:48,680
it's a question of imagining such a world as we've never been a part of. 

362
00:24:48,680 --> 00:24:54,280
The earliest evidence palaeontologists had was Eusthenopteron, 

363
00:24:54,280 --> 00:24:58,520
a fish with the precursors of our major limb bones, 

364
00:24:58,520 --> 00:25:01,520
dating from 380 million years ago. 

365
00:25:01,520 --> 00:25:04,720
But the next complete fossil specimen came after 

366
00:25:04,720 --> 00:25:06,960
a whopping gap in the record. 

367
00:25:06,960 --> 00:25:09,680
We have an animal called Eryops, 

368
00:25:09,680 --> 00:25:13,520
which is perhaps 80 million years later, 

369
00:25:13,520 --> 00:25:19,240
which has got perfectly robust limbs with fingers and toes, so those two 

370
00:25:19,240 --> 00:25:25,120
animals between them formed the sort of icons of the transition, 

371
00:25:25,120 --> 00:25:27,320
with the fish at one end of the spectrum, 

372
00:25:27,320 --> 00:25:29,760
and the tetrapod at the other end. 

373
00:25:29,760 --> 00:25:35,520
A major anatomical transformation had taken place during this time. 

374
00:25:35,520 --> 00:25:37,520
In the absence of hard evidence, 

375
00:25:37,520 --> 00:25:41,880
palaeontologists settled on the theory that periods of drought 

376
00:25:41,880 --> 00:25:44,040
had driven fish with proto-limbs onto land. 

377
00:25:45,560 --> 00:25:50,920
It was very much seen that an animal like Eusthenopteron, 

378
00:25:50,920 --> 00:25:54,760
while hopping from one drying pool to another, like a mud skipper, 

379
00:25:54,760 --> 00:25:58,920
would actually enhance its capabilities on land, 

380
00:25:58,920 --> 00:26:01,880
and that legs evolved from the fins 

381
00:26:01,880 --> 00:26:05,920
while it was on land, trying to do this hop, skip and a jump. 

382
00:26:05,920 --> 00:26:09,320
This is a hypothesis that kind of captured the public imagination, 

383
00:26:09,320 --> 00:26:14,640
and it really had a big influence on people's thinking for several decades from about the 1950s. 

384
00:26:14,640 --> 00:26:19,760
The only thing that could establish what actually happened was evidence. 

385
00:26:19,760 --> 00:26:22,680
Fossils of a creature from the middle of this gap, 

386
00:26:22,680 --> 00:26:25,840
which could reveal how the transition took place, 

387
00:26:25,840 --> 00:26:28,480
but there was one candidate for the role. 

388
00:26:29,880 --> 00:26:35,640
The only thing between Eusthenopteron and Carboniferous tetrapods with legs 

389
00:26:35,640 --> 00:26:39,640
was, for many decades, Ichthyostega. 

390
00:26:39,640 --> 00:26:42,000
Literally, that was all. 

391
00:26:44,120 --> 00:26:50,400
Ichthyostega was an intriguing tetrapod from 360 million years ago, 

392
00:26:50,400 --> 00:26:53,520
right in the middle of the fossil gap. 

393
00:26:53,520 --> 00:26:57,120
Its anatomy seemed to nod to the fish that came before 

394
00:26:57,120 --> 00:26:59,240
and the land animals that came later. 

395
00:27:00,360 --> 00:27:04,440
But the only specimen in existence was frustratingly off-limits. 

396
00:27:08,880 --> 00:27:15,240
Ichthyostega was in the hands of an eminent Swedish researcher, Erik Jarvik, 

397
00:27:15,240 --> 00:27:17,880
who had unearthed a cache of well-preserved fossils 

398
00:27:17,880 --> 00:27:21,120
on a pioneering expedition to Greenland in the 1930s. 

399
00:27:22,880 --> 00:27:25,640
In those early days, 

400
00:27:25,640 --> 00:27:29,240
these expeditions were really quite heroic, and it's a remarkable thing 

401
00:27:29,240 --> 00:27:31,760
that they brought back the material they were able to. 

402
00:27:31,760 --> 00:27:35,520
Conditions were really much more, how can I put it, primitive than today. 

403
00:27:37,360 --> 00:27:39,360
On his return from Greenland, 

404
00:27:39,360 --> 00:27:41,480
Jarvik set about a painstaking 

405
00:27:41,480 --> 00:27:43,800
investigation of this new discovery. 

406
00:28:01,280 --> 00:28:05,400
Jarvik was very hard-working and very single-minded, 

407
00:28:05,400 --> 00:28:09,800
and he was also quite talented, but not immensely talented. 

408
00:28:09,800 --> 00:28:14,640
Many things he got right, quite a few things he got wrong, 

409
00:28:14,640 --> 00:28:17,120
and he would never change his mind. 

410
00:28:17,120 --> 00:28:20,680
With Ichthyostega, Jarvik struggled, 

411
00:28:20,680 --> 00:28:25,800
and there were parts of the anatomy that he was never able to make sense of, 

412
00:28:25,800 --> 00:28:28,480
and he published it very, very slowly. 

413
00:28:36,800 --> 00:28:40,160
But the academic convention of the time meant that if other 

414
00:28:40,160 --> 00:28:44,680
researchers wanted to study this animal, they'd simply have to wait. 

415
00:28:46,360 --> 00:28:49,880
Because Jarvik had charge of the material, 

416
00:28:49,880 --> 00:28:52,840
it was felt to be his territory. 

417
00:28:52,840 --> 00:28:56,160
The etiquette of the time was really quite distinctive. 

418
00:28:56,160 --> 00:29:00,120
If somebody's in the middle of working actively on something, 

419
00:29:00,120 --> 00:29:03,360
in particular if it is stuff that they have collected themselves, 

420
00:29:03,360 --> 00:29:04,840
you don't just muscle in. 

421
00:29:04,840 --> 00:29:06,840
That would be considered very poor form. 

422
00:29:06,840 --> 00:29:10,880
The Stockholm crew had complete sovereign rights over 

423
00:29:10,880 --> 00:29:15,000
not only the material they had collected, 

424
00:29:15,000 --> 00:29:18,760
but really, the late Devonian of Greenland as a concept. 

425
00:29:20,240 --> 00:29:23,320
Other researchers' response to Jarvik's initial findings 

426
00:29:23,320 --> 00:29:25,280
only added to the delay. 

427
00:29:25,280 --> 00:29:31,160
When Jarvik published his work, British and American workers ridiculed it. 

428
00:29:31,160 --> 00:29:33,160
He was a sensitive chap, 

429
00:29:33,160 --> 00:29:36,160
and the result was a lot of bottled-up anger. 

430
00:29:36,160 --> 00:29:41,120
And it coloured his view of scientists outside Sweden 

431
00:29:41,120 --> 00:29:43,640
for the rest of his career. 

432
00:29:43,640 --> 00:29:48,520
And at this point, he felt disinclined to satisfy them 

433
00:29:48,520 --> 00:29:51,120
by publishing any more about it. 

434
00:29:51,120 --> 00:29:53,560
And he took the view that they could now wait 

435
00:29:53,560 --> 00:29:58,840
and he would work very slowly for several decades. 

436
00:29:58,840 --> 00:30:03,400
And he finally published his monograph in 1995, 

437
00:30:03,400 --> 00:30:06,080
having outlived most of his first generation of critics. 

438
00:30:06,080 --> 00:30:12,200
Jarvik's long silence left the field effectively closed. 

439
00:30:12,200 --> 00:30:14,880
This was immense frustration, 

440
00:30:14,880 --> 00:30:18,840
because the problem is that you can't really work on this kind of thing 

441
00:30:18,840 --> 00:30:20,200
without the material. 

442
00:30:20,200 --> 00:30:23,760
And if people aren't prepared to lend it to you, 

443
00:30:23,760 --> 00:30:26,320
because they are "working on it", 

444
00:30:26,320 --> 00:30:29,440
there's very little you can do about it. 

445
00:30:29,440 --> 00:30:33,200
He would frequently sit at conferences and not speak to anyone. 

446
00:30:33,200 --> 00:30:36,720
And he showed very little wish to engage in conversation. 

447
00:30:36,720 --> 00:30:41,240
Somebody did try to suggest that he should pass it on. 

448
00:30:41,240 --> 00:30:44,280
In fact, to Alec Panchen. 

449
00:30:44,280 --> 00:30:48,440
But this caused an immense rift between Jarvik and Panchen. 

450
00:30:48,440 --> 00:30:53,320
Not that Panchen felt it. But Jarvik did. 

451
00:30:53,320 --> 00:30:56,480
And that scuppered anybody else's chances 

452
00:30:56,480 --> 00:30:59,200
of getting their hands on it as well. 

453
00:30:59,200 --> 00:31:05,000
So you had this huge gap between a fish at 380 million years 

454
00:31:05,000 --> 00:31:10,200
and fairly well-known land animals at about 330-300 million years. 

455
00:31:10,200 --> 00:31:13,120
And in between, you had Ichthyostega 

456
00:31:13,120 --> 00:31:15,960
which was not very well-known at all. 

457
00:31:15,960 --> 00:31:20,160
It meant that nobody else could really think about it. 

458
00:31:20,160 --> 00:31:22,440
It was out of bounds. 

459
00:31:22,440 --> 00:31:27,800
And the subject remained moribund for decades. 

460
00:31:27,800 --> 00:31:33,200
Jenny knew that the only way to overcome the stifling etiquette 

461
00:31:33,200 --> 00:31:35,920
would be to somehow find different samples. 

462
00:31:35,920 --> 00:31:38,720
But Greenland was one of the only places in the world 

463
00:31:38,720 --> 00:31:41,840
with the right kind of Devonian rock. 

464
00:31:41,840 --> 00:31:45,920
It would be a very good area for an ambitious palaeontologist 

465
00:31:45,920 --> 00:31:48,760
to get into, IF you can find the specimens. 

466
00:31:48,760 --> 00:31:54,560
And the fact that Ichthyostega was known only from Greenland 

467
00:31:54,560 --> 00:31:58,240
which meant, at the very least, an expensive expedition, 

468
00:31:58,240 --> 00:32:00,200
meant that for many people 

469
00:32:00,200 --> 00:32:04,680
this was probably too onerous and complicated to envisage. 

470
00:32:09,600 --> 00:32:11,920
Then Jenny had an extraordinary breakthrough. 

471
00:32:11,920 --> 00:32:15,080
Brooding about Greenland, 

472
00:32:15,080 --> 00:32:17,280
she decided to ask the geologists 

473
00:32:17,280 --> 00:32:21,120
at the Earth Sciences Department over the road 

474
00:32:21,120 --> 00:32:22,760
about any trips they'd made there. 

475
00:32:22,760 --> 00:32:27,000
One recalled that a student had been to Jarvik's area of Greenland 

476
00:32:27,000 --> 00:32:32,400
in the early '70s, and had left some specimens in their storeroom. 

477
00:32:32,400 --> 00:32:34,560
In notes from a student, it said, 

478
00:32:34,560 --> 00:32:37,560
"We found tetrapods at three localities." 

479
00:32:40,080 --> 00:32:42,880
And that triggered an alarm. 

480
00:32:42,880 --> 00:32:45,320
Where is this material? 

481
00:32:45,320 --> 00:32:49,480
Jenny went down to the basement to see if she could find the fossils 

482
00:32:49,480 --> 00:32:51,920
that had been of little interest to the geologist. 

483
00:32:51,920 --> 00:32:55,960
And he pulled out a drawer full of this material. 

484
00:32:55,960 --> 00:32:59,200
He had no idea what it was he'd found. 

485
00:32:59,200 --> 00:33:04,840
It turns out that what the student had found was actually material, 

486
00:33:04,840 --> 00:33:09,480
not of Ichthyostega, but of a second animal 

487
00:33:09,480 --> 00:33:11,200
called Acanthostega, 

488
00:33:11,200 --> 00:33:15,120
which hitherto had only been known from two fragmentary specimens. 

489
00:33:15,120 --> 00:33:19,560
And what the student had found was a block 

490
00:33:19,560 --> 00:33:25,160
with three skulls of this animal together in a row. 

491
00:33:25,160 --> 00:33:29,080
I was blown away. Absolutely blown away. 

492
00:33:29,080 --> 00:33:31,360
I thought, "Oh, gosh!" 

493
00:33:31,360 --> 00:33:37,520
She was tremendously excited, because suddenly this 

494
00:33:37,520 --> 00:33:44,280
represented an opportunity to study a completely different animal. 

495
00:33:44,280 --> 00:33:48,840
A contemporary of Ichthyostega's, but obviously just from the skull 

496
00:33:48,840 --> 00:33:51,600
you could see that it was a very different animal. 

497
00:33:51,600 --> 00:33:56,360
And so it had to tell us lots of interesting, new information 

498
00:33:56,360 --> 00:33:59,960
about the evolution of tetrapods from fish. 

499
00:34:01,960 --> 00:34:03,440
Keen to know more, 

500
00:34:03,440 --> 00:34:07,360
Jenny managed to track down the student's field notebook. 

501
00:34:07,360 --> 00:34:11,040
In his scrappy notes 

502
00:34:11,040 --> 00:34:16,680
were altitudes and mountains, 

503
00:34:16,680 --> 00:34:20,160
and exactly how he'd found this material. 

504
00:34:20,160 --> 00:34:24,600
So we then knew pretty much where to go back and find this. 

505
00:34:24,600 --> 00:34:29,040
So, I mean, the obvious thing to do was try and get back there. 

506
00:34:29,040 --> 00:34:31,160
And go to the same locality. 

507
00:34:31,160 --> 00:34:35,360
The future rolled out in front of me. 

508
00:34:35,360 --> 00:34:38,400
I saw...I saw the future. 

509
00:34:38,400 --> 00:34:42,880
What was likely to happen. 

510
00:34:42,880 --> 00:34:45,920
It was astonishing that the only collection 

511
00:34:45,920 --> 00:34:51,480
of other Devonian tetrapod material was about 200-300 yards 

512
00:34:51,480 --> 00:34:54,120
from where she'd been working, and she didn't know about it. 

513
00:34:54,120 --> 00:34:56,560
None of us did. 

514
00:34:56,560 --> 00:35:01,720
That's luck, I'm afraid, by anybody's standard. 

515
00:35:01,720 --> 00:35:05,440
Well, luck and timing play into it in a big way. 

516
00:35:05,440 --> 00:35:09,320
But I think the key component, in a sense, one of the key components, 

517
00:35:09,320 --> 00:35:13,680
is spotting the chances when they come up, and leaping for them. 

518
00:35:13,680 --> 00:35:19,280
The next move was to get in touch with people in Copenhagen... 

519
00:35:20,680 --> 00:35:25,680
..and try to get them to agree to organise a joint expedition. 

520
00:35:25,680 --> 00:35:27,800
To go to Greenland, 

521
00:35:27,800 --> 00:35:31,080
Jenny would need permission from the Danish authorities. 

522
00:35:31,080 --> 00:35:33,320
Something no-one had ever previously managed, 

523
00:35:33,320 --> 00:35:37,720
owing to academic sensitivities. 

524
00:35:37,720 --> 00:35:39,840
They were always denied access. 

525
00:35:39,840 --> 00:35:44,000
This was because people in the museum in Copenhagen 

526
00:35:44,000 --> 00:35:46,400
didn't want to upset Jarvik. 

527
00:35:46,400 --> 00:35:51,440
So there's quite a lot of political activity behind this, as well. 

528
00:35:51,440 --> 00:35:54,160
Jenny saw that she had only one option. 

529
00:35:54,160 --> 00:35:56,720
To go and meet Jarvik, face to face. 

530
00:35:56,720 --> 00:36:00,680
Jenny and I went to a congress in Prague in 1985. 

531
00:36:00,680 --> 00:36:01,880
He was there. 

532
00:36:01,880 --> 00:36:05,840
She was, as I say, not a very forceful person 

533
00:36:05,840 --> 00:36:08,360
in initial conversation. 

534
00:36:08,360 --> 00:36:11,480
She's quite reserved, she's quite careful. 

535
00:36:11,480 --> 00:36:14,680
Where others had failed, Jenny's calm approach 

536
00:36:14,680 --> 00:36:18,920
and willingness to search for a fossil other than Ichthyostega 

537
00:36:18,920 --> 00:36:22,000
apparently won Jarvik round. 

538
00:36:22,000 --> 00:36:25,120
Although the conversation may have been initially a bit difficult, 

539
00:36:25,120 --> 00:36:27,440
I suspect Jarvik recognised in some ways 

540
00:36:27,440 --> 00:36:29,680
a slightly kindred personality type. 

541
00:36:29,680 --> 00:36:34,000
And not a sort of person who would go around ridiculing him 

542
00:36:34,000 --> 00:36:36,560
or in any way threatening him. 

543
00:36:36,560 --> 00:36:38,520
And in that sense, 

544
00:36:38,520 --> 00:36:41,560
I think she managed to charm him in a subtle sort of way. 

545
00:36:41,560 --> 00:36:43,800
And he was fine. 

546
00:36:43,800 --> 00:36:47,480
And that being the case, 

547
00:36:47,480 --> 00:36:52,920
the people in Copenhagen were very willing to come on board 

548
00:36:52,920 --> 00:36:58,560
and help get an expedition together to go back to Greenland. 

549
00:37:03,400 --> 00:37:09,560
Jenny now faced organising the most challenging fossil hunt of her life. 

550
00:37:09,560 --> 00:37:11,080
Enlisting her husband Rob, 

551
00:37:11,080 --> 00:37:14,920
and her PhD student, Per Ahlberg, to help her. 

552
00:37:14,920 --> 00:37:18,600
We'd never done anything like this before. I was actually terrified. 

553
00:37:18,600 --> 00:37:20,240
I had no idea really what to expect, 

554
00:37:20,240 --> 00:37:22,240
except that it's going to be difficult. 

555
00:37:22,240 --> 00:37:27,600
We were going in helicopters, and everybody knows helicopters crashed. 

556
00:37:27,600 --> 00:37:29,440
And polar bears eat you. 

557
00:37:29,440 --> 00:37:32,000
We knew this was going to be a hard trip. 

558
00:37:32,000 --> 00:37:34,680
We knew that we would be out in the field, 

559
00:37:34,680 --> 00:37:36,480
camping 300km inside the Arctic Circle. 

560
00:37:36,480 --> 00:37:38,560
We knew it was going to be cold. 

561
00:37:38,560 --> 00:37:41,200
We knew it was going to be hard work, climbing up mountains. 

562
00:37:41,200 --> 00:37:42,800
And we knew we were not very fit. 

563
00:37:42,800 --> 00:37:45,800
My girlfriend knitted me a nice, warm, new jumper. 

564
00:37:45,800 --> 00:37:48,600
I've still got it, too. It's got fish on it. 

565
00:37:48,600 --> 00:37:51,720
I've still got the same girlfriend, too. 

566
00:37:51,720 --> 00:37:57,640
In July 1987, a Danish support team deposited Jenny, Per and Rob 

567
00:37:57,640 --> 00:38:01,360
on the side of Stensio Bjerg, in the remote northeast of Greenland. 

568
00:38:06,960 --> 00:38:08,800
Now suddenly, here you are. 

569
00:38:08,800 --> 00:38:12,200
There's nobody for 100 miles in any direction. 

570
00:38:13,600 --> 00:38:15,560
That is isolated. 

571
00:38:15,560 --> 00:38:18,880
Having set up camp and made things work on that level, 

572
00:38:18,880 --> 00:38:21,440
we now had to go out and find something. 

573
00:38:21,440 --> 00:38:24,720
And if we didn't find anything of any significance, 

574
00:38:24,720 --> 00:38:26,600
it would be rather embarrassing 

575
00:38:26,600 --> 00:38:29,240
and rather a lot of money spent for nothing. 

576
00:38:32,320 --> 00:38:36,400
Looking for fossils can be a thankless task. 

577
00:38:36,400 --> 00:38:39,640
Because most of the time, you don't find anything. 

578
00:38:44,000 --> 00:38:47,080
For the first few days, we couldn't find where we were supposed to be. 

579
00:38:47,080 --> 00:38:49,440
We weren't even sure we were on the right mountain. 

580
00:38:49,440 --> 00:38:50,720
And we thought, 

581
00:38:50,720 --> 00:38:53,440
"We're here for 6 weeks and not going to go back with anything?" 

582
00:38:53,440 --> 00:38:59,200
We were walking on very steep scree slope, 

583
00:38:59,200 --> 00:39:01,720
made up of small slivers of rock. 

584
00:39:01,720 --> 00:39:04,800
Very loose, very insecure. 

585
00:39:04,800 --> 00:39:08,680
And at the bottom, we could see what appeared to be a sheer cliff 

586
00:39:08,680 --> 00:39:10,960
waiting for us to fall over it. 

587
00:39:10,960 --> 00:39:12,280
So we gave up. 

588
00:39:12,280 --> 00:39:15,080
And the three of us trudged back down the mountain, 

589
00:39:15,080 --> 00:39:18,320
feeling very depressed and ashamed, as you can imagine. 

590
00:39:18,320 --> 00:39:21,280
Jenny, as I recall, was getting quite worried about 

591
00:39:21,280 --> 00:39:25,720
whether anything useful was going to come out of this. 

592
00:39:25,720 --> 00:39:32,080
Because we were looking for white bone in darkish brown rock. 

593
00:39:32,080 --> 00:39:38,000
So anything white caught our eyes and we thought for an instant, 

594
00:39:38,000 --> 00:39:40,200
"I've found a fossil." 

595
00:39:40,200 --> 00:39:43,520
But a lot of the time, it was bird shit. 

596
00:39:46,120 --> 00:39:49,680
Almost one week in, the exhausted party 

597
00:39:49,680 --> 00:39:52,240
launched a third attempt on the mountain, 

598
00:39:52,240 --> 00:39:54,640
approaching it at a different level. 

599
00:39:54,640 --> 00:39:56,680
One of us, I forget who now, 

600
00:39:56,680 --> 00:40:00,560
picked up a little slab with a piece of bone on it, 

601
00:40:00,560 --> 00:40:02,800
showed it to Jenny, who gave a kind of whoop, 

602
00:40:02,800 --> 00:40:07,480
because this was part of the back of the skull of Acanthostega. 

603
00:40:07,480 --> 00:40:10,000
It was a piece of the same fossil animal 

604
00:40:10,000 --> 00:40:13,800
that Jenny had found in the drawer in Cambridge. 

605
00:40:13,800 --> 00:40:16,480
She realised that they must be close to the area 

606
00:40:16,480 --> 00:40:18,840
described in the student's notes. 

607
00:40:18,840 --> 00:40:21,480
As we started hacking at the rock, 

608
00:40:21,480 --> 00:40:24,800
hack a block off, break it up... 

609
00:40:24,800 --> 00:40:26,680
another skull. 

610
00:40:26,680 --> 00:40:30,480
We realised we are in this big kind of apron of fallen material 

611
00:40:30,480 --> 00:40:32,120
from some locality higher up. 

612
00:40:32,120 --> 00:40:35,160
We kept going up and it kept getting richer and richer, 

613
00:40:35,160 --> 00:40:38,840
and I found a lower jaw actually disappearing into the rock. 

614
00:40:38,840 --> 00:40:43,160
So there we were, and we had found the locality. 

615
00:40:43,160 --> 00:40:45,400
Now, of course, once we'd done that, 

616
00:40:45,400 --> 00:40:48,120
then we were really in business. 

617
00:40:48,120 --> 00:40:51,760
The team had discovered a whole strata of rock 

618
00:40:51,760 --> 00:40:54,240
full of Acanthostega remains. 

619
00:40:54,240 --> 00:40:58,400
And it looked like it contained more than just scattered bones. 

620
00:41:01,200 --> 00:41:04,000
The dream of the vertebrate palaeontologist 

621
00:41:04,000 --> 00:41:06,840
is to find a complete, articulated specimen, 

622
00:41:06,840 --> 00:41:10,520
with every bone in place, so you tip it out the rock and you can see what's going on. 

623
00:41:10,520 --> 00:41:13,200
One of the specimens that we found, 

624
00:41:13,200 --> 00:41:16,040
it was clear that it was a head at one end 

625
00:41:16,040 --> 00:41:18,640
and then leading back from there, 

626
00:41:18,640 --> 00:41:24,040
in sort of a glancing sunlight, was a row of bumps, 

627
00:41:24,040 --> 00:41:27,280
which looked as though they might be vertebrae. 

628
00:41:27,280 --> 00:41:30,400
We didn't know what we'd found. 

629
00:41:30,400 --> 00:41:32,280
But we knew it was going to be exciting. 

630
00:41:32,280 --> 00:41:37,000
It looked as if they had found one of palaeontology's holy grails. 

631
00:41:37,000 --> 00:41:38,760
The first complete specimen 

632
00:41:38,760 --> 00:41:41,480
of an early Devonian tetrapod. 

633
00:41:41,480 --> 00:41:44,800
This was it. This was the key, really, 

634
00:41:44,800 --> 00:41:47,720
to the rest of my career. 

635
00:41:47,720 --> 00:41:51,520
The discovery of those materials was probably the most exciting thing 

636
00:41:51,520 --> 00:41:53,520
after falling in love. 

637
00:41:56,520 --> 00:42:02,320
The party packed a metric tonne of fossils into crates 

638
00:42:02,320 --> 00:42:06,640
and returned to Cambridge. 

639
00:42:14,000 --> 00:42:17,480
DRILLING 

640
00:42:17,480 --> 00:42:21,440
Once home, they faced the monumental task 

641
00:42:21,440 --> 00:42:25,480
of bringing the rocks to life. 

642
00:42:25,480 --> 00:42:27,480
Here, you are dealing with an animal 

643
00:42:27,480 --> 00:42:29,480
and you don't know what it's going to look like. 

644
00:42:29,480 --> 00:42:32,320
Nobody knows what it's going to look like. 

645
00:42:32,320 --> 00:42:34,840
And you are able to piece together for the first time 

646
00:42:34,840 --> 00:42:37,960
a style of creature that nobody perhaps has previously seen. 

647
00:42:37,960 --> 00:42:41,600
Trying to imagine what these animals were like in life 

648
00:42:41,600 --> 00:42:45,320
is really what it's about for me. 

649
00:42:45,320 --> 00:42:48,560
I kind of wish we had a time machine. 

650
00:42:48,560 --> 00:42:51,680
That we could go back and look at the animals. 

651
00:42:51,680 --> 00:42:54,040
Because a lot of us are zoologists at heart. 

652
00:42:54,040 --> 00:42:56,680
These are animals that happen to be in the rocks. 

653
00:42:56,680 --> 00:43:00,800
Jenny now assembled another team 

654
00:43:00,800 --> 00:43:05,640
to undertake the painstaking work of fossil preparation. 

655
00:43:05,640 --> 00:43:07,640
When I started working on that material 

656
00:43:07,640 --> 00:43:10,800
using the adapted dental equipment that we use, 

657
00:43:10,800 --> 00:43:13,280
picks and drills and so forth, 

658
00:43:13,280 --> 00:43:16,760
I realised this was going to, er... 

659
00:43:16,760 --> 00:43:19,040
This was going to take a hell of a long time. 

660
00:43:19,040 --> 00:43:24,040
The Greenland rock is very hard, so it takes a long time to get 

661
00:43:24,040 --> 00:43:27,280
a very small amount of material out of the specimen. 

662
00:43:27,280 --> 00:43:29,680
You had these long periods of relatively 

663
00:43:29,680 --> 00:43:31,160
methodical preparation, 

664
00:43:31,160 --> 00:43:34,600
punctuated by great periods of excitement, so... 

665
00:43:34,600 --> 00:43:37,360
You're obsessive in doing this work, 

666
00:43:37,360 --> 00:43:40,200
because you don't know what you're going to find next. 

667
00:43:40,200 --> 00:43:41,360
And it's absorbing. 

668
00:43:41,360 --> 00:43:44,000
It can be nerve-wracking 

669
00:43:44,000 --> 00:43:49,440
because you don't really know where the bone is. 

670
00:43:49,440 --> 00:43:55,400
You have to be extremely careful and take it down bit by bit. 

671
00:43:55,400 --> 00:43:57,880
It's all a matter of colour and texture, 

672
00:43:57,880 --> 00:44:00,480
and being able to see subtle differences 

673
00:44:00,480 --> 00:44:03,400
as you expose different levels. 

674
00:44:05,440 --> 00:44:09,720
Finding out whether the Acanthostega fossils had legs was critical. 

675
00:44:09,720 --> 00:44:14,800
They could be the key to understanding how limbs had evolved. 

676
00:44:17,080 --> 00:44:21,240
I think we were pretty open-minded about it in the first place 

677
00:44:21,240 --> 00:44:25,560
because for all we knew, it could have had a fin instead of a limb. 

678
00:44:25,560 --> 00:44:29,160
I mean, other features of the anatomy suggested 

679
00:44:29,160 --> 00:44:31,960
that it really was quite primitive. 

680
00:44:31,960 --> 00:44:36,640
Jenny gave Mike Coates the job of preparing the body 

681
00:44:36,640 --> 00:44:40,320
of the most complete-looking specimen. 

682
00:44:40,320 --> 00:44:42,960
How much of the animal was there, we weren't sure. 

683
00:44:42,960 --> 00:44:46,200
So, of course, part of the rest of that was unpacking blocks. 

684
00:44:46,200 --> 00:44:48,480
Again, sitting in dusty museum basements 

685
00:44:48,480 --> 00:44:51,080
just trying to fit these things back together again, 

686
00:44:51,080 --> 00:44:53,840
how they'd been in the field before they'd been broken up. 

687
00:44:53,840 --> 00:45:01,160
He started work on what we thought was going to be a humerus arm bone. 

688
00:45:01,160 --> 00:45:04,680
And, indeed, that's what it turned out to be. 

689
00:45:04,680 --> 00:45:08,840
And it was pretty comparable with the humerus of a tetrapod. 

690
00:45:08,840 --> 00:45:11,760
The upper arm suggested 

691
00:45:11,760 --> 00:45:15,000
that this must be a land-walking amphibian. 

692
00:45:15,000 --> 00:45:18,560
But to their surprise, when Mike uncovered the lower arm bones, 

693
00:45:18,560 --> 00:45:22,160
these resembled those of Eusthenopteron, the fish ancestor. 

694
00:45:22,160 --> 00:45:25,640
It was an astonishing mix. 

695
00:45:25,640 --> 00:45:29,520
So then he started working on the adjoining block, 

696
00:45:29,520 --> 00:45:31,560
starting from the edge and working in. 

697
00:45:31,560 --> 00:45:35,200
And the first thing he found is a digit. 

698
00:45:35,200 --> 00:45:42,040
A row of elements joined one to the other, like fingers. 

699
00:45:42,040 --> 00:45:43,280
So he found a finger. 

700
00:45:43,280 --> 00:45:48,280
Digits meant a hand or a foot, not present in Eusthenopteron. 

701
00:45:48,280 --> 00:45:50,040
But that wasn't all. 

702
00:45:50,040 --> 00:45:53,680
So he went a bit further, found another one. 

703
00:45:54,760 --> 00:45:56,240
And another one. 

704
00:45:56,240 --> 00:45:59,240
And another one. And another one. 

705
00:46:00,720 --> 00:46:03,720
And he thought, "Ah, I've found five. 

706
00:46:03,720 --> 00:46:06,240
"Shall I carry on, or shall I just leave it?" 

707
00:46:06,240 --> 00:46:10,080
So he thought, "Just for the sake of completeness, I'll do a bit more." 

708
00:46:10,080 --> 00:46:14,280
And he went round and eventually came up with eight of them. 

709
00:46:14,280 --> 00:46:17,080
And that was... 

710
00:46:17,080 --> 00:46:19,520
when the fun really started. 

711
00:46:19,520 --> 00:46:22,120
The assumption had always been 

712
00:46:22,120 --> 00:46:26,000
that tetrapods had evolved with five digits. 

713
00:46:26,000 --> 00:46:29,760
It made a huge impact, because it seemed so outlandish. 

714
00:46:29,760 --> 00:46:33,240
And there were one or two people who wondered initially 

715
00:46:33,240 --> 00:46:35,800
whether what they had found was simply the two limbs 

716
00:46:35,800 --> 00:46:37,240
lying on top of each other, 

717
00:46:37,240 --> 00:46:39,880
giving that effect. Understandably enough. 

718
00:46:39,880 --> 00:46:42,840
But very quickly, when a few people had seen the materials, 

719
00:46:42,840 --> 00:46:46,680
it was acknowledged that this was in fact exactly what they said it was. 

720
00:46:46,680 --> 00:46:51,080
There had been a big debate for many decades about how digits had arisen 

721
00:46:51,080 --> 00:46:54,080
and why they look the way they do. 

722
00:46:54,080 --> 00:46:56,920
And the eight-digited limb of Acanthostega 

723
00:46:56,920 --> 00:46:59,000
landed kind of smack in this. 

724
00:47:00,840 --> 00:47:02,240
Jenny began to wonder 

725
00:47:02,240 --> 00:47:06,440
whether she needed to rethink the most basic assumptions. 

726
00:47:06,440 --> 00:47:11,520
I think as Mike was preparing the forelimb, 

727
00:47:11,520 --> 00:47:14,320
it just became obvious 

728
00:47:14,320 --> 00:47:16,520
that the forearms stretched out like that. 

729
00:47:16,520 --> 00:47:18,240
Really couldn't bend very far. 

730
00:47:18,240 --> 00:47:22,160
They just do not look like weight-bearing limbs. 

731
00:47:22,160 --> 00:47:27,080
So we began to think of what else it could do. 

732
00:47:27,080 --> 00:47:30,200
What was it being used for? 

733
00:47:30,200 --> 00:47:31,680
It was a confusing picture. 

734
00:47:31,680 --> 00:47:33,200
The arms didn't bend in a way 

735
00:47:33,200 --> 00:47:35,920
that would allow Acanthostega to walk. 

736
00:47:35,920 --> 00:47:37,400
And the eight-toed hand 

737
00:47:37,400 --> 00:47:41,720
looked more like a paddle than a foot it could stand on. 

738
00:47:41,720 --> 00:47:43,960
Then Jenny had a radical thought. 

739
00:47:43,960 --> 00:47:49,600
What if Acanthostega's limbs and feet hadn't evolved on land? 

740
00:47:49,600 --> 00:47:55,640
Perhaps limbs evolved before walking. Perhaps for some other purpose. 

741
00:47:55,640 --> 00:47:57,640
Swimming. 

742
00:48:00,240 --> 00:48:05,280
That idea, that limbs evolved in water first, 

743
00:48:05,280 --> 00:48:09,680
was quite revolutionary. 

744
00:48:09,680 --> 00:48:13,600
Maybe making their way through swampy, reedy, mucky streams. 

745
00:48:13,600 --> 00:48:16,400
Paddling in water, rather than walking on land. 

746
00:48:16,400 --> 00:48:19,480
In other words, something entirely different first. 

747
00:48:19,480 --> 00:48:22,000
And then only later used for walking with. 

748
00:48:22,000 --> 00:48:25,720
Just as she had with the stapes earlier in her career, 

749
00:48:25,720 --> 00:48:31,480
Jenny's creative vision was turning the received wisdom on its head. 

750
00:48:31,480 --> 00:48:35,480
But she knew she needed further proof. 

751
00:48:35,480 --> 00:48:38,200
She focused on the skull. 

752
00:48:38,200 --> 00:48:40,520
Inside the throat 

753
00:48:40,520 --> 00:48:43,360
were a series of gill bars, 

754
00:48:43,360 --> 00:48:48,680
which looked just the same as those that you find in fish. 

755
00:48:48,680 --> 00:48:51,880
So it's gill breathing. And then Mike found the tail. 

756
00:48:51,880 --> 00:48:55,320
Which had got long, long fin rays. 

757
00:48:55,320 --> 00:48:57,640
Which is useless out of water, 

758
00:48:57,640 --> 00:48:59,640
so that sort of completes the picture 

759
00:48:59,640 --> 00:49:01,960
of primitive tetrapods 

760
00:49:01,960 --> 00:49:04,280
living in water, being mainly aquatic, 

761
00:49:04,280 --> 00:49:07,080
and using their limbs for swimming. 

762
00:49:07,080 --> 00:49:08,680
Everybody expected Acanthostega 

763
00:49:08,680 --> 00:49:11,160
to look like a terrestrial animal from the neck back. 

764
00:49:11,160 --> 00:49:13,040
Nobody expected it to have gills. 

765
00:49:13,040 --> 00:49:15,440
It was a paradigm shift. 

766
00:49:15,440 --> 00:49:17,440
The decades-old textbook image 

767
00:49:17,440 --> 00:49:19,880
of a fish lumbering onto land on its fins 

768
00:49:19,880 --> 00:49:21,200
could only be wrong. 

769
00:49:24,520 --> 00:49:27,320
Other fossil evidence of plant life supported the idea 

770
00:49:27,320 --> 00:49:30,160
that these creatures lived in newly-formed marshland. 

771
00:49:32,960 --> 00:49:37,120
It seemed that the very first legs evolved not for walking, 

772
00:49:37,120 --> 00:49:42,080
but as a tactic for moving through dense vegetation in swamps. 

773
00:49:42,080 --> 00:49:46,080
Things can be evolved for one purpose 

774
00:49:46,080 --> 00:49:48,720
and then used later 

775
00:49:48,720 --> 00:49:51,600
in a slightly modified form for another thing. 

776
00:49:51,600 --> 00:49:53,440
And the idea that limbs 

777
00:49:53,440 --> 00:49:55,720
were not originally used for walking 

778
00:49:55,720 --> 00:49:57,960
might be counterintuitive. 

779
00:49:57,960 --> 00:50:01,880
But I think that's only because as humans and as terrestrial animals, 

780
00:50:01,880 --> 00:50:04,120
we kind of think walking must be 

781
00:50:04,120 --> 00:50:07,160
the be all and end all of what limbs are for. 

782
00:50:07,160 --> 00:50:08,600
But, of course, it isn't. 

783
00:50:10,240 --> 00:50:15,720
Once Jenny was ready to publish, her findings were well received. 

784
00:50:15,720 --> 00:50:17,880
I think she realised pretty quickly 

785
00:50:17,880 --> 00:50:20,360
that this was Nature-standard material. 

786
00:50:20,360 --> 00:50:22,360
That the journal, Nature, 

787
00:50:22,360 --> 00:50:26,760
which is one of the most prestigious scientific journals, 

788
00:50:26,760 --> 00:50:29,960
would actually publish this sort of stuff. 

789
00:50:29,960 --> 00:50:36,280
From when we started publishing the full descriptions of Acanthostega, 

790
00:50:36,280 --> 00:50:38,600
the head, the body, the limbs, 

791
00:50:38,600 --> 00:50:44,160
it became the model for a Devonian tetrapod. 

792
00:50:44,160 --> 00:50:50,240
Or a primitive tetrapod that could have given rise to later ones. 

793
00:50:50,240 --> 00:50:52,600
And Ichthyostega was marginalised. 

794
00:50:52,600 --> 00:50:56,840
But there was a remaining mystery about Ichthyostega. 

795
00:50:56,840 --> 00:50:59,200
Jarvik, the Swedish researcher 

796
00:50:59,200 --> 00:51:02,560
who had hung on to the only specimen for 50 years, 

797
00:51:02,560 --> 00:51:04,480
had described his animal 

798
00:51:04,480 --> 00:51:07,720
as having five toes. 

799
00:51:07,720 --> 00:51:11,280
We'd also found a hind limb of Ichthyostega 

800
00:51:11,280 --> 00:51:13,320
in the same expedition. 

801
00:51:13,320 --> 00:51:20,360
And preparation of that showed that it had seven digits on its foot. 

802
00:51:20,360 --> 00:51:26,240
So putting that together, a pattern of more than five digits 

803
00:51:26,240 --> 00:51:30,040
seemed to be what early tetrapods had. 

804
00:51:30,040 --> 00:51:32,280
If you go back and look Jarvik's specimen, 

805
00:51:32,280 --> 00:51:34,360
what he thought was one big toe, cracked, 

806
00:51:34,360 --> 00:51:37,360
with a lot of cracks in it, is actually these three toes. 

807
00:51:37,360 --> 00:51:39,480
So again, I think he was a little bit annoyed 

808
00:51:39,480 --> 00:51:41,560
that she'd interpreted it and he hadn't. 

809
00:51:41,560 --> 00:51:46,760
Jenny Clack had again seen what others had missed. 

810
00:51:46,760 --> 00:51:51,720
But Jarvik was unwilling to accept that science had moved on. 

811
00:51:51,720 --> 00:51:57,160
He finally published his definitive text on Ichthyostega. 

812
00:51:57,160 --> 00:52:01,360
And in it, he used photographic evidence of his own 

813
00:52:01,360 --> 00:52:04,080
to dismiss Clack's specimens as freaks. 

814
00:52:04,080 --> 00:52:08,320
Jarvik didn't really believe our story, 

815
00:52:08,320 --> 00:52:12,000
or our interpretation of the evidence. 

816
00:52:12,000 --> 00:52:14,400
So he considered that the material 

817
00:52:14,400 --> 00:52:16,840
of Ichthyostega and Acanthostega 

818
00:52:16,840 --> 00:52:20,200
were both mutants that we happen to have found. 

819
00:52:20,200 --> 00:52:24,680
You know, we know how rare mutants are in normal life. 

820
00:52:24,680 --> 00:52:29,360
And the idea that we had found two of them was a bit silly. 

821
00:52:29,360 --> 00:52:35,360
He just had the mindset that he had this safe knowledge 

822
00:52:35,360 --> 00:52:39,320
and that what we had found contradicted that. 

823
00:52:39,320 --> 00:52:40,720
So it was wrong. 

824
00:52:40,720 --> 00:52:42,200
Jenny's not like that at all. 

825
00:52:42,200 --> 00:52:44,760
She's very open-minded about her work. 

826
00:52:44,760 --> 00:52:48,400
One of the things that I appreciated very much being her student 

827
00:52:48,400 --> 00:52:53,560
was the sense that the student was always welcome to disagree 

828
00:52:53,560 --> 00:52:55,160
with the supervisors. 

829
00:52:55,160 --> 00:52:58,840
I don't imagine Jenny has ever thought herself to be infallible 

830
00:52:58,840 --> 00:53:00,480
in any aspect of this stuff. 

831
00:53:00,480 --> 00:53:03,680
This, of course, is the mark of a really good scientist. 

832
00:53:03,680 --> 00:53:07,160
Because that way, your science becomes self-correcting. 

833
00:53:07,160 --> 00:53:10,080
You're not going to veer off to the side and land in the ditch 

834
00:53:10,080 --> 00:53:13,720
which, in a sense, is what happened with Jarvik and his interpretations. 

835
00:53:13,720 --> 00:53:18,960
You will continue to head onwards towards a more and more 

836
00:53:18,960 --> 00:53:22,000
accurate understanding of the animals you're working with. 

837
00:53:22,000 --> 00:53:26,200
Other palaeontologists embraced Clack's theory 

838
00:53:26,200 --> 00:53:28,680
of the aquatic origin of limbs, 

839
00:53:28,680 --> 00:53:31,880
opening up a whole new area of research. 

840
00:53:31,880 --> 00:53:35,480
All of a sudden, Devonian tetrapods went from being 

841
00:53:35,480 --> 00:53:38,760
a marginal subject area to one of the hottest areas in palaeontology. 

842
00:53:38,760 --> 00:53:40,360
It's catalysed the whole thing. 

843
00:53:40,360 --> 00:53:44,320
We are now finding Devonian tetrapods worldwide. 

844
00:53:44,320 --> 00:53:46,120
And if you look at a graph 

845
00:53:46,120 --> 00:53:50,160
of the number of taxa known of Devonian tetrapods, 

846
00:53:50,160 --> 00:53:51,640
it's gone like that. 

847
00:53:51,640 --> 00:53:54,520
So it's almost exponential increase. 

848
00:53:54,520 --> 00:53:56,000
It's had a huge impact. 

849
00:53:56,000 --> 00:54:00,880
It's as though the field had become moribund, 

850
00:54:00,880 --> 00:54:04,600
and now it's been completely unlocked, and it's vibrant. 

851
00:54:04,600 --> 00:54:06,240
It was pretty radical. 

852
00:54:06,240 --> 00:54:09,640
Yeah, we rewrote the textbooks, effectively. 

853
00:54:09,640 --> 00:54:12,480
How long that will last, I'm not sure. 

854
00:54:12,480 --> 00:54:14,680
But that remains to be seen. 

855
00:54:18,640 --> 00:54:20,880
The self-deprecating girl 

856
00:54:20,880 --> 00:54:22,560
who never expected to do a PhD 

857
00:54:22,560 --> 00:54:26,480
had reached the pinnacle of her field. 

858
00:54:26,480 --> 00:54:28,920
Whoever you asked in the subject would say 

859
00:54:28,920 --> 00:54:30,280
Jenny is the world leader 

860
00:54:30,280 --> 00:54:32,520
in research on the origin of tetrapods. 

861
00:54:32,520 --> 00:54:33,880
She's the great pioneer. 

862
00:54:33,880 --> 00:54:37,360
She's the one who led the way and opened up this area 

863
00:54:37,360 --> 00:54:39,880
that we others have kind of come in to 

864
00:54:39,880 --> 00:54:42,240
and continue to contribute to. 

865
00:54:42,240 --> 00:54:46,400
And she continues to drive that subject forward to this day. 

866
00:54:46,400 --> 00:54:50,920
She is, for most of us, the leading worker in this field. 

867
00:54:50,920 --> 00:54:53,760
And as a measure of this, 

868
00:54:53,760 --> 00:54:58,400
we do talk about the Clack theory of the origin of tetrapods. 

869
00:54:58,400 --> 00:55:02,760
And there are very few other workers in vertebrate palaeontology 

870
00:55:02,760 --> 00:55:07,280
where a theory, a current theory, is immediately understood 

871
00:55:07,280 --> 00:55:11,280
by virtue of the name of the person who came up with it. 

872
00:55:11,280 --> 00:55:16,120
I think Jenny definitely deserves all the awards 

873
00:55:16,120 --> 00:55:18,040
and status that she's getting. 

874
00:55:18,040 --> 00:55:21,320
I'm sure she'd say she doesn't, but I think she does. 

875
00:55:21,320 --> 00:55:27,120
It's characteristic of a really good palaeontologist that they have, 

876
00:55:27,120 --> 00:55:30,080
how can I put it, the imagination 

877
00:55:30,080 --> 00:55:36,520
to understand these wretched remains as a one-time living organism. 

878
00:55:36,520 --> 00:55:39,360
And Jenny is very good at this. 

879
00:55:39,360 --> 00:55:45,600
Jenny became Professor Clack in 1997. 

880
00:55:45,600 --> 00:55:48,920
But her greatest accolade came when she was invited to become 

881
00:55:48,920 --> 00:55:51,960
the first woman in her field to join the Royal Society. 

882
00:55:54,760 --> 00:55:58,040
I suppose I had secretly wanted that for years 

883
00:55:58,040 --> 00:55:59,920
but never thought it would happen. 

884
00:55:59,920 --> 00:56:06,920
And I was nominated by the director of the museum at the time, 

885
00:56:06,920 --> 00:56:12,400
and I thought, "It's not going to get anywhere." 

886
00:56:12,400 --> 00:56:16,240
But it did. It was one of the most thrilling days of my life, 

887
00:56:16,240 --> 00:56:17,840
when I got that letter. 

888
00:56:17,840 --> 00:56:20,280
I don't think she made a great fuss about it. 

889
00:56:20,280 --> 00:56:23,480
I don't think she went round saying, "Hey, look at me!" 

890
00:56:23,480 --> 00:56:27,000
That's just not in her persona. 

891
00:56:28,360 --> 00:56:32,520
I was immensely proud, and I went round shouting, "Hey, look at her!" 

892
00:56:32,520 --> 00:56:38,640
But I'm still constantly amazed by how my career has gone. 

893
00:56:38,640 --> 00:56:41,880
And it's not finished yet, of course. 

894
00:56:41,880 --> 00:56:43,720
But, erm...truly astonishing. 

895
00:56:46,720 --> 00:56:50,160
Since describing Acanthostega, Professor Clack has remained 

896
00:56:50,160 --> 00:56:53,160
at the forefront of tetrapod research, 

897
00:56:53,160 --> 00:56:57,600
with over 140 publications. 

898
00:56:57,600 --> 00:57:00,400
And she continues to make new discoveries 

899
00:57:00,400 --> 00:57:02,760
that fill the gaps in the fossil record. 

900
00:57:02,760 --> 00:57:06,480
But she accepts that her theories could yet be superseded. 

901
00:57:06,480 --> 00:57:08,240
All our discoveries... 

902
00:57:08,240 --> 00:57:09,960
Fossils will remain, 

903
00:57:09,960 --> 00:57:13,400
but the interpretations can be overturned any day. 

904
00:57:13,400 --> 00:57:15,800
All this knowledge is provisional. 

905
00:57:15,800 --> 00:57:18,200
And this is something that people don't really 

906
00:57:18,200 --> 00:57:19,680
understand about science. 

907
00:57:19,680 --> 00:57:21,600
That it's not about certainty. 

908
00:57:21,600 --> 00:57:24,080
It's not necessarily even about facts. 

909
00:57:24,080 --> 00:57:27,760
It's about questions. And the answers that you give. 

910
00:57:27,760 --> 00:57:31,400
A lesson in life's impermanence, 

911
00:57:31,400 --> 00:57:34,600
that perhaps only palaeontology's long perspective 

912
00:57:34,600 --> 00:57:36,600
could have given her. 

913
00:57:36,600 --> 00:57:43,640
We should remember that we are only here temporarily. 

914
00:57:43,640 --> 00:57:47,760
We do need to bear in mind that 

915
00:57:47,760 --> 00:57:53,080
something will evolve to take our place at some point. 

916
00:57:53,080 --> 00:57:54,760
And I like to speculate about 

917
00:57:54,760 --> 00:57:57,400
which group of animals that might come from. 

918
00:57:58,840 --> 00:58:00,920
My betting is on rodents, actually. 

919
00:58:27,360 --> 00:58:31,400
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